Institute of Wildlife Management and Wildlife Biology, University of Sopron, 9400 Sopron, Hungary
10.48311/jibs.12.01.81
Abstract
A checklist with distribution data of the collembolan genus Pseudosinella of Hungary is presented in this paper. In addition to available relevant references, unpublished data from newly collected material is also included. To date, the occurrence of 23 species of this genus has been confirmed in Hungary, for which a dichotomous key is also provided. Since the publication date of the Hungarian Collembola checklist, a total of nine additional Pseudosinella species have been recorded in the country. In this study, P. edax Gisin, 1967, P. nosekiRusek, 1985, and P. thibaudi Stomp, 1977 are cited for the first time. Up to now, three species of Pseudosinella, namely P. argenteaLoksa, 1960 nec Folsom, P. csafordensisWinkler & Mateos, 2018, and P. dungeriWinkler & Mateos, 2018, are recognized as endemic to Hungary.
Arbea, J.I. (2013) A new cave species of Pseudosinella (Collembola, Entomobryomorpha, Entomobryidae) from Sima del Campamento (Jaén, Southern Iberian Peninsula). Arquivos Entomolóxicos, 8, 303–308.
Arbea, J.I. & Jordana, R. (1990) New species of Pseudosinella and Lepidocyrtus from Navarra (Northern Iberian Peninsula). Spixiana, 13, 25–31.
Bellinger, P.F., Christiansen, K.A. & Janssens, F. (1996–2025) Checklist of the Collembola of the World. http://www.collembola.org [Accessed October 15, 2025]
Bogojević, J. (1971) Prilog poznavanju faune Collembola Deliblatske peščare. Matica Srpska Journal for Natural Sciences, 40, 153–156. [in Serbian with English summary]
Börner, F. (1903) Über neue Altweltliche Collembolen, nebst Bemerkungen zur Systematik der Isotominen und Entomobryinen. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1903,129–182.[in German]https://doi.org/10.5962/bhl.part.29866
Buşmachiu, G. (2010) Checklist of springtails (Collembola) from the Republic of Moldova. Travaux Du Muséum National d’Histoire Naturelle “Grigore Antipa”, 53 (1), 149–160. https://doi.org/10.2478/v10191-010-0011-x
Buşmachiu, G. (2021) Fauna Colembolelor (Hexapoda: Collembola) din Republica Moldova. Căpățină Print, Chișinău. 200 pp. [in Romanian]
Christian, E. (1987) Catalogus Faunae Austriae, Teil XIIa.: U.-Kl.: Collembola (Springschwänze). Verlag der Österreichischen Akademie der Wissenschaften, Wien. 83 pp. [in German]
Christian, E. (1997) Die wirbellosen Tiere der Hermannshöhle bei Kirchberg am Wechsel, Niederösterreich. Die Höhle, 50, 205–224. [in German]
Christiansen, K.A. & Bellinger, P.F. (1998) The Collembola of North America north of the Rio Grande; A taxonomic analysis. 2nd Edition. Grinnell College, Grinnell, IA. 1518 pp.
Christiansen, K.A., Bellinger, P. & da Gama, M.M. (1990) Computer Assisted Identification of Specimens of Pseudosinella (Collembola: Entomobryidae). Revue d’Écologie et de Biologie du Sol, 27 (2), 231–246.
Cipola, N.G., Oliveira, J.V.L.C., Bellini, B.C., Ferreira, A.S., Lima, E.C.A., Brito, R.A., Stievano, L.C., Souza, P.G.C. & Zeppelini, D. (2020) Review of eyeless Pseudosinella Schäffer (Collembola, Entomobryidae, and Lepidocyrtinae) from Brazilian caves. Insects, 11 (3), 194. https://doi.org/10.3390/insects11030194
Cuchta, P. & Shrubovych, J. (2015) Soil Collembola communities in montane coniferous forests of the Bohemian Forest. Suva Gabreta, 21 (2), 149–156.
Dallai, R. & Malatesta, E. (1982) Ricerche sui Collemboli. XXVI. Collemboli cavernicoli italiani. Lavori della Società Italiana di Biogeografia, 7, 173–194. [in Italian with English summary]
Dányi, L. (2011) Cave dwelling springtails (Collembola) of Hungary: a review. Soil Organisms, 83 (3), 419–432.
Dányi, L. & Traser, Gy. (2007) An annotated checklist of the springtail fauna of Hungary (Hexapoda: Collembola). Opuscula Zoologica, 38, 3–82.
Deharveng, L. & Thibaud, J.M. (1989) Acquisitions recentes sur les Insectes Collemboles cavernicoles d’Europe. Mémoires de Biospéologie, 16, 145–151. [in French with English summary]
Denis, J.R. (1931) Sur la faune Italienne des Collemboles, IV. Memorie della Societa Entomologica Italiana, 10, 80–85. [in French]
Dudich, E. (1930) Az Aggteleki barlang állatvilágának élelemforrásai. Állattani Közlemények, 27, 62–85. [in Hungarian with German summary]
Dudich, E. (1932) Biologie der Aggteleker Tropfsteinhöhle „Baradla” in Ungarn. Verlag Speläologisches Institut, Wien, 13, 1–246. [in German]
Dunger, W. (1975) Collembolen aus dem Börzsöny–Gebirge. Folia Historico-Naturalia Musei Matraensis, 3, 11–33. [in German with English summary]
Farkas, J. (1995) Collembola fauna from the shore of Lake Balaton, Hungary. Opuscula Zoologica, 27–28, 45–47.
Farkas, J., Szathmáry, K. & Strenner, A. (1998) Ecofaunistical investigations of Collembola, Araneae and Coleoptera in mosaic-like habitats in the Cinege Valley, Hungary. Opuscula Zoologica, 31, 49–61.
Fiera, C. (2013) Species of the genus Pseudosinella (Collembola: Entomobryidae) in the Romanian fauna. Acta Oecologica Carpatica, 6, 53–66.
Fjellberg, A. (2007) The Collembola of Fennoscandia and Denmark, Part II: Entomobryomorpha and Symphypleona. Fauna Entomologica Scandinavica, 42, 1–264.
Flórián, N., Ladányi, M., Ittzés, A., Kröel-Dulay, G., Ónodi, G., Mucsi, M., Szili-Kovács, T., Gergócs, V., Dányi, L. & Dombos, M. (2019) Effects of single and repeated drought on soil microarthropods in a semi-arid ecosystem depend more on timing and duration than drought severity. PLoS ONE, 14 (7), e0219975. https://doi.org/10.1371/journal.pone.0219975
Fratello, B., Sabatini, M.A., Mola, L., Uscidda, C. & Gessa, C. (1989) Effects of agricultural practices on soil arthropoda: Organic and mineral fertilizers in alfalfa fields. Agriculture, Ecosystems & Environment, 27 (1–4), 227–239.https://doi.org/10.1016/0167-8809(89)90088-1
Gergócs, V., Flórián, N., Tóth, Z., Szili-Kovács, T., Mucsi, M. & Dombos, M. (2022) Crop species and year affect soil-dwelling Collembola and Acari more strongly than fertilisation regime in an arable field. Applied Soil Ecology, 173,104390. https://doi.org/10.1016/j.apsoil.2022.104390
Gisin, H. (1955) Recherches sur la relation entre la faune endogée de Collemboles et les qualités agri-coliques de sols viticoles. Revue suisse de Zoologie, 62, 601–648. [in French]
Gisin, H. (1967) Espèces nouvelles et lignées évolutives de Pseudosinella endogés (Collembola). Memórias e Estudos do Museu Zoologico da Universidade de Coimbra, 301, 1–25. [in French]
Gisin, H. & Gama, M.M. da (1969) Espèces nouvelles de Pseudosinella cavernicoles (Insecta: Collembola). Revue Suisse de Zoologie, 76, 143–181.[in French] https://doi.org/10.5962/bhl.part.146030
Gruia, M. (1970) Donnés sur certains Collemboles lapidicoles et lithoclasicoles des Carpates. Travaux de l’Institut “Emile Racovitza”, 9, 191–195. [in French]
Harta, I., Winkler, D. & Füleki, G. (2018) Impact of former long-term fertilization on springtail communities in a reforested experimental area. Columella – Journal of Agricultural and Environmental Sciences, 5 (1),13–25.https://doi.org/10.18380/szie.colum.2018.5.1.13
Harta, I., Simon, B., Vinogradov, S. & Winkler, D. (2021) Collembola communities and soil conditions in forest plantations established in an intensively managed agricultural area. Journal of Forestry Research, 32, 1819–1832.https://doi.org/10.1007/s11676-020-01238-z
Jordana, R. (2012) Capbryinae and Entomobryini. In: Dunger, W. & Burkhardt, U. (eds) Synopses on Palaearctic Collembola. Soil Organisms, 84 (1), 1–391.
Jordana, R. & Baquero, E. (2005) New species of Pseudosinella (Collembola, Entomobryidae) from Tesla Mountains, Burgos, Spain. Serie Zoologica, 29 (1), 1–7.
Kampmann, T. (1987) Untersuchungen an Collembolenzönosen in Wäldern der Bundesrepublik Deutschland. Ph.D. Dissertation, University of Bonn. 179 pp.
Kaprus, I.J. & Goblyk, K. (2015) Ecological and sozological evaluation of soils of the Transcarpathian Lowland with using Collembola communities. Proceedings of the State Natural History Museum, 31, 45–58.
Kopeszki, H. & Trockner, V. (1994) Effects of skiing on the collembolan fauna of an alpine meadow in Grodental (South Tyrol). Zoologischer Anzeiger, 233, 221–239.
Koledin, D., Matic, R., Tancic, N. & Stamenkovic, S. (1988) Collembola community dynamics in beech forest habitats on Jastrebac mountain (Yugoslavia). Arhiv bioloskih Nauka, 40 (1–4), 53–68.
Kovács, I.E. (1953) Biológiai terepszemle az aggteleki Békebarlangban. In: Jakucs, L. (ed.) A Békebalang Felfedezése. Művelt Nép, Budapest, pp. 86–91.
Kovács, I.E. (1954) Első vizsgálatok az aggteleki Békebarlang faunáján. Állattani Közlemények,44(1–2), 95–98. [in Hungarian with German summary]
Kováč, Ľ. (2000) A review of the distribution of cave Collembola (Hexapoda) in the Western Carpathians. Mémoires de Biospéologie, 27, 71–76.
Kováč, L. & Rusek, J. (2012) Redescription of two troglobiotic species of the genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) from the Western Carpathians. Zootaxa, 3341, 32–45. https://doi.org/10.11646/zootaxa.3341.1.2
Kováč, Ĺ., Žurovcová, M., Raschmanová, N., Jureková, N., Papáč, V. & Parimuchová, A. (2023) Troglomorphic adaptations on the northern European frontier: the phylogeny of the cave Pseudosinella (Hexapoda, Collembola) in the Western Carpathians. Frontiers in Ecology and Evolution,11, 1169911.https://doi.org/10.3389/fevo.2023.1169911
Kseneman, M. (1935) Apterygota z území rybníku Lednickych II. Zvláštní otisk z Časopisu Národního Musea, 1931, 1–27. [in Czech with French summary]
Kühnelt, W. (1963) Über den Einfluβ des Mycels von Clitocybe infundibuliformis auf Streufauna. In: Doeksen, J. & van der Drift, J. (eds) Soil Organisms. North Holland Pub. Co., Amsterdam, pp. 281–287.
Loksa, I. (1956a) Zoologische Untersuchungen von Kollembolen im Bükkgebirge. Acta Zoologica Academiae Scientiarum Hungaricae, 2 (4), 379–419. [in German]
Loksa, I. (1956b) Die zönologische Untersuchung von Kollembolen in einer ungarischen Querceto- Potentilletum Albae-Assoziation. Acta Zoologica Academiae Scientiarum Hungaricae, 2, 199–243. [in German]
Loksa, I. (1960) Faunistisch–systematische und ökologische Untersuchungen in der Lóczy-Höhle bei Balatonfüred (Biospeologica Hungarica, XI.). Annales Universitas Scientiarum Budapestinensis, Sectio Biologica, 3, 253–266. [in German]
Loksa, I. (1961) Ökologisch-faunistischetischeUntersuchungen in der Freiheitshöhle bei Égerszög. (Biospeologica Hungarica, XIII.). Acta Zoologica Academiae Scientiarum Hungaricae, 7, 219–230. [in German]
Loksa, I. (1962) Über die Landarthropoden der István-, Forrás- und Szeleta-Höhle bei Lillafüred. Karszt– és Barlangkutatás, 3, 59–81. [in German]
Loksa, I. (1966) Die bodenzoozönologischen Verhältnisse der Flaumeichen–Buschwälder Südostmittel-europas. Akadémiai Kiadó, Budapest. 437 pp. [in German]
Loksa, I. (1977) Quantitative Analysen des Collembolenbesatzes im Jahre 1975. im Untersuchungsgebiet bei Síkfőkút. Annales Universitatis Scientiarum Budapestinensis de Rolando Eötvös, Nominate, Sectio Biologica, 18–19, 205–218. [in German]
Loksa, I. (1978a) Die Collembolen-Fauna der Urwacholder aus der Umgebung von Barcs. Dunántúli Dolgozatok, Természettudományi Sorozat, 1, 51–64. [in German]
Loksa, I. (1978b) Mikrohabitate und ihre Bedeutung für die Verteilung der Collenbolengemeinschaften in einem Hainbuchen-Eichenbestand. Opuscula Zoologica, 15 (1–2),93–117. [in German with English summary]
Loksa, I. (1980) A siroki Nyírjes-tó ugróvillás rovarai (Collembola). Folia Historico–Naturalia Musei Matraensis, 6, 107–114.[in Hungarian]
Loksa, I. (1991) The Collembola fauna of the Bátorliget Nature Reserves (NE Hungary). In: Mahunka, S. (ed.) The Bátorliget Nature Reserves – after forty years. Studia Naturalia, 1 (1), 269–277.
Loksa, I. & Loksa, I. (1996a) Adatok a Tétényi-fennsík ízeltlábú faunájához. In: Simon, T. (ed.) Természeti kincsek Dél-Budán; A Tétényi-fennsík és a Háros-sziget növény- és állatvilága, természetvédelme. Cserépfalvi Kiadó és Zöld Jövő Környezetvédelmi Egyesület, Budapest, pp. 40–51. [in Hungarian]
Loksa, I. & Loksa, I. (1996b) Adatok a Háros-sziget ízeltlábú faunájához. In: Simon, T. (ed.) Természeti kincsek Dél-Budán; A Tétényi-fennsík és a Háros-sziget növény- és állatvilága, természetvédelme. Cserépfalvi Kiadó és Zöld Jövő Környezetvédelmi Egyesület, Budapest, pp. 70–79. [in Hungarian]
Loksa, I. & Rubio, I. (1966) Anagben zu den Kenntnissen über die Collembolenfauna des Bakony- Gebirges. Opuscula Zoologica,6 (1), 139–156. [in German]
Mehrafroz Mayvan, M., Sadeghi-Namaghi, H., Shayanmehr, M. & Greenslade, P. (2022) Contribution to the knowledge of Entomobryomorpha (Hexapoda: Collembola) from Northeastern Iran with new records and a key to the species. Journal of Insect Biodiversity and Systematics, 8 (2), 151–174. https://doi.org/10.52547/jibs.8.2.151
Mock, A., Šašková, T., Raschmanová, N., Jászay, T., Ľuptáčik, P., Rendoš, M., Tajovský, K. & Jászayová, A. (2015) An introductory study of subterranean communities of invertebrates in forested talus habitats in southern Slovakia. Acta Societatis Zoologicae Bohemicae, 79, 237–250.
Paclt, J. (1960) Über eine kleine Apterygoten-Ausbeute aus Ungarn. Opuscula Zoologica, 3 (3–4), 155–158. [in German]
Palkó, Á., Ónodi, G., Rédei, T. & Winkler, D. (2020) Talajfaunisztikai- és ökológiai vizsgálatok alföldi reliktum homoki tölgyesekben és a helyükön létesített idegenhonos faállományokban. Erdészettudományi Közlemények, 10 (1–2), 123–137.[in Hungarian with English summary] http://doi.org/10.17164/EK.2020.010
Ponge, J.P. (2000) Acidophilic Collembola: living fossils? Contributions from the Biological Laboratory, Kyoto University, 29, 65–74.
Rusek, J. (1979) Three new Pseudosinella species from Czechoslovakia (Collembola, Entomobryidae). Acta Entomologica Bohemoslovaca, 76, 255–265.
Rusek, J. (1985) New Palearctic Lepidocyrtus and Pseudosinella species (Collembola: Entomobryidae). Vestnik Ceskoslovenske Spolecnosti Zoologicke, 49, 132–146.
Shayanmehr, M., Yoosefi Lafooraki, E., Tully, T., D’Haese, C. & Ghajar Sepanlou, M. (2024) New records of springtails (Hexapoda: Collembola) for Iran from the Bula Hyrcanian forest. Journal of Insect Biodiversity and Systematics, 10 (1), 31–42. https://doi.org/10.61186/jibs.10.1.31
Sławska, M., Bruckner, A. & Sławski, M. (2017) Edaphic Collembola assemblages of European temperate primeval forests gradually change along a forest-type gradient. European Journal of Soil Biology, 80, 92–101.https://doi.org/10.1016/j.ejsobi.2017.05.003
Stach, J. (1929) Verzeichnis der Apterygogenea Ungarns. Annales Historico-Naturales Musei Nationalis Hungarici, 26, 269–312.[in German]
Sterzyńska, M. (1995) Collembola of secondary succession of the pine forests of Puszcza Białowieska. Fragmenta Faunistica, 37, 354–369.
Stomp, N. (1977) Nouvelle contribution a l’etude des Pseudosinella endoges d’Europe (Collemboles, Entomobryidae). Revue d’Écologie et de Biologie du Sol, 14 (1),231–239.
Stomp, N. (1986) Le groupe de Pseudosinella vandeli (Insectes, Collemboles), dans les Alpes Orientales. In: Dallai, R. (ed) Second International Seminar on Apterygota. University of Siena, Siena, pp. 85–91. [in French]
Szent-Ivány, J. (1940) Adatok a Kőszeg-vidéki ugróvillás rovarok (Collembola) ismeretéhez. Dunántúli Szemle, 7 (7–8), 425–439. [in Hungarian]
Strouhal, H. & Vornatscher, J. (1975) Katalog der rezenten Höhlentiere Österreichs. Annales Naturhistorisches Museum Wien, 79, 401–542. [in German]
Szigeti, N., Berki, I., Vityi, A. & Winkler, D. (2022) Soil mesofauna and herbaceous vegetation patterns in an agroforestry landscape. Agroforestry Systems, 96 (4), 773–786. https://doi.org/10.1007/s10457-022-00739-6
Tschubby (2025) Reliefkarte Ungarn.https://w.wiki/GHie [Accessed October 15, 2025]
Traser, Gy. (1980) Adatok a farkasgyepűi bükkösök avarszintjének Collembola (ugróvillás) faunájához. Erdészeti és Faipari Tudományos Közlemények, 2, 19–23.[in Hungarian with English summary]
Traser, Gy. (1981) Talajinszekticidek hatása a talajfaunára. Erdészeti és Faipari Tudományos Közlemények,3(2), 129–138. [in Hungarian with English summary]
Traser, Gy. (1995) The Collembola fauna of Őrség (Insecta: Collembola). In: Víg, K. (ed.) Natural History of Őrség Landscape conservation Area I. Savaria Múzeum, Szombathely, pp. 43–47.
Traser, Gy. (1997) Egy szigetközi nemesnyáras biodiverzitása: a talajfauna. In: Pájer, J. & Fabich, E. (eds) V. Erdészeti Szakmai Konferencia kiadványa a WoodTech keretében. Sopron, pp. 57–64. [in Hungarian]
Traser, Gy. (1999) Springtails of the Aggtelek National Park (Hexapoda: Collembola). In: Mahunka, S. (ed.) The fauna of the Aggtelek National Park. Akadémiai Kiadó, Budapest, pp. 49–59.
Traser, Gy. (2002a) The Collembola of the Fertő–Hanság National Park. In: Mahunka, S. (ed.) The fauna of the Fertő-Hanság National Park. MTM, Budapest, pp. 259–270.
Traser, Gy. (2002b) Borókás–nyáras erdők ugróvillás (Insecta: Collembola) faunája. In: Barna, T. (ed.) Alföldi Erdőkért Egyesület Kutatói nap 2001-2002., Alföldi Erdőkért Egyesület, Gyula, pp. 46–52.
Traser Gy. (2003) Hansági nemesnyár és éger erdők ugróvillás (Insecta: Collembola) faunája. III. Kárpát-medencei Biológiai Szimpózium kiadványa. Magyar Biológiai Társaság, Budapest, pp. 153–157. [in Hungarian]
Traser, Gy. (2006) A gyarmatpusztai vadgesztenyés ugróvillás (Insecta: Collembola) faunájának diverzitása. Folia Musei Historico-Naturalis Bakonyiensis, 23, 33–39. [in Hungarian with English summary]
Traser, Gy. (2008) Rovartani érdekességek (Insecta: Collembola) egy patak menti égererdőből. In: Barna, T. (ed.) Alföldi Erdőkért Egyesület Kutatói Nap 2008. Alföldi Erdőkért Egyesület, Szeged, pp. 120–125. [in Hungarian with English summary]
Traser, Gy. (2010) A Collembola fauna (Hexapoda: Entognatha) Porva körzetében a harmadik Biodiverzitás Nap gyűjtései alapján. A Bakonyi Természettudományi Múzeum Közleményei, 27, 49–54.
Traser, Gy. & Horváth-Szováti, E. (2004) Tájhasznosítás hatása a talajfaunára: ugróvillás (Parainsecta: Collembola) közösségek diverzitása Csévharaszton. In: Barna, T. (ed.) Alföldi Erdőkért Egyesület Kutatói Nap 2004. Alföldi Erdőkért Egyesület, Kecskemét, pp. 51–62. [in Hungarian with English summary]
Traser, Gy. & Horváth-Szováti, E. (2006) Land–use intensity and diversity parameters: Collembola (Insecta) communities in Csévharaszt (Hungary). In: Csóka, Gy., Hirka, A. & Koltay, A. (eds) Biotic Damage in Forests. Proceedings of the IUFRO (WP7.03.10) Symposium held in Mátrafüred, Hungary, pp. 329–339.
Traser, Gy., Szűcs, P. & Winkler, D. (2006) Collembola diversity of moss habitats in the Sopron Region, NW-Hungary. Acta Silvatica et Lignaria Hungarica, 2, 69–80.
Traser, Gy., Winkler, D. & Molnár, M. (2011) A mezőföldi Szent László-víz völgyének ugróvillás (Collembola) faunája. A Bakonyi Természettudományi Múzeum Közleményei, 28, 71–79. [in Hungarian with English summary]
Varga, Z. (1995) Geographical patterns of biological diversity in the Palearctic region and the Carpathian Basin. Acta Zoologica Academiae Scientiarum Hungaricae, 41 (2), 71–92.
Vedovini, A. (1968) Contribution a l'inventaire faunistique des cavites souterraines des environs de Marseille. Annales de Speleologie, 23, 229–242.[in French]
Winkler, D. (2014) Collembolan response to red mud pollution in Western Hungary. Applied Soil Ecology, 83, 219–229.https://doi.org/10.1016/j.apsoil.2013.07.006
Winkler, D. (2019) Ugróvillások (Collembola). In: Kárpáti, L. (ed.) Soproni Tájvédelmi Körzet - Monografikus tanulmányok a Soproni-hegység természeti és kulturális értékeiről. Szaktudás Kiadó Ház Zrt., Budapest, pp. 133–135.[in Hungarian]
Winkler, D. & Mateos, E. (2018) New species of Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) from Hungary. Zootaxa, 4382 (2), 347–366. https://doi.org/10.11646/zootaxa.4382.2.7
Winkler, D. & Tóth, V. (2012) Effects of afforestation with pines on Collembola diversity in the limestone hills of Szárhalom (West Hungary). Acta Silvatica et Lignaria Hungarica, 8, 9–20. https://doi.org/10.2478/v10303-012-0001-8
Winkler, D. & Traser, Gy. (2012a) Collembola Diversity in Agricultural Environments (Lajta Project, Western Hungary). In: Proceedings of the International Scientific Conference on Sustainable Development and Ecological Footprint. 26–27 March 2012, Sopron, Hungary, pp. 1–5.
Winkler, D. & Traser, Gy. (2012b) Eco-faunistic study on the Collembola fauna in the Vasvár-Nagymákfa area. Natura Somogyiensis, 22, 39–52. https://doi.org/10.24394/NatSom.2012.22.39
Winkler, D., Korda, M. & Traser, Gy. (2012) Rare and little known Collembola species from Hungary. Opuscula Zoologica,43 (1), 89–95.
Winkler, D., Bidló, A., Bolodár-Varga, B., Erdő, Á. & Horváth, A. (2018) Long-term ecological effects of the red mud disaster in Hungary: Regeneration of red mud flooded areas in a contaminated industrial region. Science of the Total Environment, 644, 1292–1303. https://doi.org/10.1016/j.scitotenv.2018.07.059
Winkler, D., Németh, M.T. & Fiera, C. (2021) A new species of Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) from Hungary and Romania, with redescription of the related species Pseudosinella huetheri Stomp, 1971. ZooKeys, 1063, 121–137. https://doi.org/10.3897/zookeys.1063.73094
Articles in Press, Corrected Proof Available Online from 12 December 2025
Winkler,D. (2025). The genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) in Hungary. (e27964). Journal of Insect Biodiversity and Systematics, (), e27964 doi: 10.48311/jibs.12.01.81
MLA
Winkler,D. . "The genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) in Hungary" .e27964 , Journal of Insect Biodiversity and Systematics, , , 2025, e27964. doi: 10.48311/jibs.12.01.81
HARVARD
Winkler D. (2025). 'The genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) in Hungary', Journal of Insect Biodiversity and Systematics, (), e27964. doi: 10.48311/jibs.12.01.81
CHICAGO
D. Winkler, "The genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) in Hungary," Journal of Insect Biodiversity and Systematics, (2025): e27964, doi: 10.48311/jibs.12.01.81
VANCOUVER
Winkler D. The genus Pseudosinella Schäffer, 1897 (Collembola, Entomobryidae) in Hungary. J. Insect Biodivers. Syst., 2025; (): e27964. doi: 10.48311/jibs.12.01.81
